...in their swift, short steps

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Senior Member
Could someone, please, advise me on this preposition "IN" "These waders or shorebirds live in marshes, at the edge of lagoons and in muddy, slimy lands. They can be seen in intertidal areas pecking about in the mud, actively running around in their swift, short steps, searching for food, fundamentally worms, prawns, molluscs and dead fish" Thanks in advance
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