reference of "who"

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Blue Apple

Senior Member
Persian (Iran)
Does the highlighted "who" refer to "Marsden Hartley and Charles Demuth" or "Jasper Johns and Andy Warhol"? I think the latter, am I right?

Context:
Kenneth Silver argues that key works by the US artists Jasper Johns and Andy Warhol of the late 1950s and early 1960s “began to label something—male homosexuality—that had hitherto been considered too unworthy or too dangerous to name.” Earlier twentieth-century American artists had also dealt with gay culture— Silver discusses Marsden Hartley and Charles Demuth as important precedents for later artists who took on homosexual life and culture as a subject, and another art historian, Richard Meyer, points to the painter Paul Cadmus (Art and Politics by Claudia Mesch).
 
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